Hejnol Group - Selected Publications


Research Articles

Reviews and Book Chapters

  • Hejnol A, Vellutini BC (2017) Larval Evolution: I'll tail you later… Curr Biol, 29, R21-R23

  • Hejnol A, Pang K (2016). Xenacoelomorpha’s significance for understanding bilaterian evolution Curr Opin Dev Gen 39, 48-54

  • Hejnol A, Dunn CW (2016). Animal Evolution: Are Phyla Real? Curr. Biol., 26, R424-R426

  • Arnone MI, Hejnol A (2015). Genomics going wild: Marine sampling for studies of evolution and development Mar Genomics, doi:10.1016/j.margen.2015.11.003

  • Hejnol A, Lowe CJ (2015). Embracing the comparative approach: how robust phylogenies and broader developmental sampling impacts the understanding of nervous system evolution Phil Trans R Soc B 370: 20150045 - link

  • Adell T, Martín-Durán JM, Salo E, Cebria F (2015). Platyhelminthes. Evolutionary Developmental Biology of Invertebrates Volume 2, ed. Wanninger A, Springer Verlag, pp. 21-40.

  • Hejnol A, Rentzsch F (2015). Primer: Neural Nets. Curr. Biol. 25:R782-R786 - link.

  • Hejnol A (2015). Acoelomorpha. Structure and Evolution of Invertebrate Nervous Systems eds. Schmidt-Rhaesa A, Hartzsch S, Purschke G, Oxford University Press, Oxford, pp. 56-61.

  • Hejnol A (2015). Acoelomorpha and Xenoturbellida. Evolutionary Developmental Biology of Invertebrates Volume 1, ed. Wanninger A., Springer Verlag, pp. 203-14.

  • Hejnol A (2015). Cycloneuralia. Evolutionary Developmental Biology of Invertebrates Volume 3, ed. Wanninger A, Springer Verlag, pp. 1-13.

  • Hejnol A (2015). Gnathifera. Evolutionary Developmental Biology of Invertebrates Volume 2, ed. Wanninger A, Springer Verlag, pp. 1-12.

  • Hejnol A (2015). Gastrotricha. Evolutionary Developmental Biology of Invertebrates Volume 2, ed. Wanninger A, Springer Verlag, pp. 13-19.

  • Hejnol A, Martín-Durán JM (2015). Getting to the bottom of anal evolution. Zool. Anz., doi: 10.1016/j.jcz.2015.02.006 - link

  • Hejnol A, Lowe C (2014). Animal Evolution: Stiff or Squishy Notochord Origins? Curr. Biol. 24, R1131-R1133

  • Dunn CW, Giribet G, Edgecombe GD, Hejnol A (2014). Animal Phylogeny and Its Evolutionary Implications. Annu. Rev. Ecol. Evol. Syst. 45:371-395

  • Hejnol A (2014). News and Views: Evolutionary Biology: Excitation about jelly nerves. Nature, 510, doi:10.1038/nature13340, Nature Podcast Not so Simple

  • Martín-Durán JM, Hejnol A (2014). Priapulida (penis worms): Implications of embryonic development. 306-309 In: McGraw-Hill 2014 Yearbook of Science & Technology, McGraw-Hill Education, New York

  • Hejnol A (2012). News and Views: Evolutionary Biology: Muscle's dual origins. Nature, 487, 181-182.

  • Martín-Durán JM, Egger B (2012). Developmental diversity in free-living flatworms. EvoDevo, 3:7 - abstract

  • Edgecombe GD, Giribet G, Dunn CW, Hejnol A, Kristensen RM, Neves RC, Rouse GW, Worsaae K, Sørensen MV. (2011) Higher-level metazoan relationships: recent progress and remaining questions. Org. Divers. Evol., Volume 11, Number 2, 151-172.

  • Hejnol A (2011). Mark Q. Martindale: Shedding new light on developmental diversity Int. J. Dev. Biol., 55: 237-242.

  • Hejnol A (2010). A Twist in Time--The Evolution of Spiral Cleavage in the Light of Animal Phylogeny Integr. Comp. Biol.; 50, 695-706, doi: 10.1093/icb/icq103

Publications prior to arrival at the Sars Centre

  • Hejnol A, Obst M, Stamatakis A, Ott M, Rouse GW, Edgecombe GD, Martinez P, Baguñà J, Bailly X, Jondelius U, Wiens M, Müller, WEG, Seaver E, Wheeler WC, Martindale MQ, Giribet G, Dunn CW (2009). Assessing the root of bilaterian animals with scalable phylogenomic methods. Proc R Soc B, 276, 4261-4270.


  • Hejnol A and Martindale MQ (2009). The mouth, the anus and the blastopore - open questions about questionable openings. Animal Evolution: Genomes, Fossils and Trees, eds. Littlewood D.T.J. & Telford, M.J., Oxford University Press, pp. 33-40. (pdf)


  • Martindale MQ and Hejnol A (2009). A developmental perspective: changes in the position of the blastopore during bilaterian evolution. Dev Cell, 17, 162-174.


  • Bourlat SJ and Hejnol A (2009). Quick Guide: Acoels. Curr. Biol., 19, R279-R280.


  • Hejnol A and Martindale MQ (2009). The mouth, the anus and the blastopore - open questions about questionable openings. In: “Animal Evolution: Genomes, Fossils and Trees”, eds. Littlewood D.T.J. & Telford, M.J., Oxford University Press, pp. 33-40.


  • Giribet G, Dunn CW, Edgecombe GD, Hejnol A, Martindale MQ, Rouse GW (2009). Assembling the Spiralian Tree of Life. In: “Animal Evolution: Genomes, Fossils and Trees”, eds. Littlewood D.T.J. & Telford, M.J., Oxford University Press, pp. 52-64.


  • Hejnol A and Martindale MQ (2008). Acoel development indicates the independent evolution of the bilaterian mouth and anus. Nature, 456, 382-386. (listen to the nature podcast)


  • Dunn CW, Hejnol A, Matus DQ, Pang K, Browne WE, Smith SA, Seaver E, Rouse GW, Obst M, Edgecombe GD, Sørensen MV, Haddock SHD, Schmidt-Rhaesa A, Okuso A, Kristensen RM, Wheeler WC, Martindale MQ, Giribet G (2008). Broad phylogenomic sampling improves resolution of the Animal Tree of Life. Nature, 452, 745-749.


  • Hejnol A and Martindale MQ (2008). Acoel development supports a simple planula-like urbilaterian. Phil Trans R Soc B, 363, 1493-1501.


  • Henry JQ, Hejnol A, Perry KJ, Martindale MQ (2007). Homology of ciliary bands in spiralian trochophores. Integ Comp Biol, 47, 865-871.


  • Hejnol A, Martindale MQ, Henry JQ (2007). High-resolution fate map of the snail Crepidula fornicata: the origins of ciliary bands, nervous system, and muscular elements. Dev Biol, 305, 63-76.


  • Hejnol A, Schnabel R, Scholtz G (2006). A 4D-microscopic analysis of the germ band in Porcellio scaber (Malacostraca, Peracarida) – developmental and phylogenetic implications. Dev Genes Evol, 216, 755-767.


  • Matus DQ, Copley RR, Dunn CW, Hejnol A, Eccleston H, Halanych KH, Martindale MQ, Telford MJ (2006). Broad taxon and gene sampling indicate that chaetognaths are protostomes. Curr Biol, 16, R575-R576.


  • Schnabel R, Bischoff M, Hintze A, Schulz A-K, Hejnol A, Meinhardt H, Hutter H (2006). Global cell sorting in the C. elegans embryo defines a new mechanism for pattern formation. Dev Biol, 294, 418-431.


  • Hejnol A and Schnabel R (2006). What a couple of dimensions can do for you: Comparative developmental studies using 4D-microscopy – examples from tardigrade development. Integ Comp Biol, 46, 151–161.


  • Hejnol A and Schnabel R (2005). The eutardigrade Thulinia stephaniae has an indeterminate development and the potential to regulate early blastomere ablations. Development, 132, 1349-1361.


  • Hejnol, A., Scholtz, G. (2004) Clonal analysis of Distal-less and engrailed expression patterns during early morphogenesis of uniramous and biramous crustacean limbs. Dev Genes Evol 214, 473-485.


  • Hejnol, A., Pfannenstiel, H.-D. (1998) Myosin and actin are necessary for polar lobe formation and resorption in Ilyanassa obsoleta embryos. Dev Genes Evol 208, 229-233.



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